B. holtonis var. holtonis

Memoirs of the New York Botanical Garden. New York: The Garden, 1900- v. 8 1952-1954: Page: 36-40 http://www.biodiversitylibrary.org/item/150964 Junction of the Provinces of Guayas, Cafiar, Chimborazo and Bolivar: plants much-branched, climbing to several meters, stem and pedicels red, leaves pale green with red margin, flowers flushed with pink; foothills of the western cor­diller: near the village of Bucay, 1,000-1,250 ft., E-3662.
Pests Diseases

• Classification: Species
• Plant Type: Shrub-like
• Botanical Section: Lepsia
• Origin: America, Colombia, Ecuador
• Originator: A. de Candolle, Ann. Sci. nat., Bot. ser. 4.
• Date of Origin: 1859